Humankind is responsible for conditioning the distribution of species, including the African elephant. From its great physical form to its fascinating social behaviour, Loxodonta africana can trigger an admiration in some and be undervalued by others, often ending up being a victim of human activities. Increasingly, the apparent impossibility of the coexistence of these two species represents a problem for the conservation of the large African mammal. Here it was reviewed the published bibliography about the African savannah elephant ( L. africana ), between July 2015 and December 2019. The Web of Science (WoS) database enabled the search for the words “ Loxo donta african a ”. From this research, we obtained 310 articles, 149 of them were analysed. Elephants have characteristics that define its uniqueness that are constant ly reported, such as: its cognition and extraordinary memory, factors that influence its movements and its distribution, its specific vocalization and the strong dependence on social connections. Threats to the conservation of the species and the urgency of interventions are also an important subject of discussion. Poaching for ivory represents a major obstacle on conservation and has been drastically reducing the number of elephant populations and causi ng future consequences for those who thrive. Law enforcement in r educing/prohibit poaching has proven to be insufficient, so there is an urgent need to combat the demand for ivory. Also, it is necessary to bypass the conflict, promote tolerance and enable cohabitation. We believe that educating human populations to better understand the species may be the difference to guaran tee a future for the species. Conservationists may be the enablers of these ed ucational actions, however, to make the difference, it is necessary to know the species in detail. Mastering knowledge in these behavioural areas will contribute to improve mitigation strategies, which could make the difference to guarantee a future for the species.
Blanc, J. (2008) Loxodonta africana. 2008 IUCN Red List of Threatened Species. Gland, Switzerland, 8235. http://www.iucnredlist.org
Petracca, L.S., Funston, P.J., Henschel, P., Cohen, J.B., Maclennan, S. and Frair, J.L. (2019) Modeling Community Occupancy from Line Transect Data: A Case Study with Large Mammals in Post-War Angola. Animal Conservation, 1-14. https://doi.org/10.1111/acv.12555
Schlossberg, S., Chase, M.J. and Griffin, C.R. (2018) Poaching and Human Encroachment Reverse Recovery of African Savannah Elephants in South-East Angola Despite 14 Years of Peace. PLoS ONE, 13, e0193469. https://doi.org/10.1371/journal.pone.0193469
Stalmans, M.E., Massad, T.J., Peel, M.J.S., Tarnita, C.E. and Pringle, R.M. (2019) War-Induced Collapse and Asymmetric Recovery of Large-Mammal Populations in Gorongosa National Park, Mozambique. PLoS ONE, 14, e0212864. https://doi.org/10.1371/journal.pone.0212864
Bastille-Rousseau, G., Wall, J., Douglas-Hamilton, I., Lesowapir, B., Loloju, B., Mwangi, N. and Wittemyer, G. (2019) Landscape-Scale Habitat Response of African Elephants Shows Strong Selection for Foraging Opportunities in a Human Dominated Ecosystem. Ecography, 43, 149-160. https://doi.org/10.1111/ecog.04240
Branco, P.S., Merkle, J.A., Pringle, R.M., King, L., Tindall, T., Stalmans, M. and Long, R.A. (2019) An Experimental Test of Community-Based Strategies for Mitigating Human-Wildlife Conflict around Protected Areas. Conservation Letters, 13, e12679. https://doi.org/10.1111/conl.12679
Mpakairi, K.S., Ndaimani, H., Kuvawoga, P.T. and Madiri, H.T. (2019) Human Settlement Drives African Elephant (Loxodonta africana) Movement in the Sebungwe Region, Zimbabwe. African Journal of Ecology, 57, 531-538. https://doi.org/10.1111/aje.12639
Gaynor, K.M., Branco, P.S., Long, R.A., Goncalves, D.D., Granli, P.K. and Poole, J.H. (2018) Effects of Human Settlement and Roads on Diel Activity Patterns of Elephants (Loxodonta africana). African Journal of Ecology, 56, 872-881. https://doi.org/10.1111/aje.12552
Youldon, D.A., Abell, J., Briffitt, J.S., Chama, L., Channings, M.D., Kilundo, A., Larsen, C.K., Sakala, D. and Schulte, B.A. (2017) Patch-Occupancy Survey of Elephant (Loxodonta africana) Surrounding Livingstone, Zambia. Koedoe, 59, a1372. https://doi.org/10.4102/koedoe.v59i1.1372
Buchholtz, E., Fitzgerald, L., Songhurst, A., McCulloch, G. and Stronza, A. (2019) Overlapping Landscape Utilization by Elephants and People in the Western Okavango Panhandle: Implications for Conflict and Conservation. Landscape Ecology, 34, 1411-1423. https://doi.org/10.1007/s10980-019-00856-1
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Mitigation Strategies
Poaching
Wildlife Conservation
Cook, R.M., Henley, M.D. and Parrini, F. (2015) Elephant Movement Patterns in Relation to Human Inhabitants in and around the Great Limpopo Transfrontier Park. Koedoe, 57, a1298. https://doi.org/10.4102/koedoe.v57i1.1298
Smit, J., Pozo, R.A., Cusack, J.J., Nowak, K. and Jones, T. (2019) Using Camera Traps to Study the Age-Sex Structure and Behaviour of Crop-Using Elephants Loxodonta africana in Udzungwa Mountains National Park, Tanzania. Oryx, 53, 368-376. https://doi.org/10.1017/S0030605317000345
Branco, P.S., Merkle, J.A., Pringle, R.M., Pansu, J., Potter, A.B., Reynolds, A., Stalmans, M. and Long, R.A. (2019) Determinants of Elephant Foraging Behaviour in a Coupled Human-Natural System: Is Brown the New Green? Journal of Animal Ecology, 88, 780-792. https://doi.org/10.1111/1365-2656.12971
Matseketsa, G., Muboko, N., Gandiwa, E., Kombora, D.M. and Chibememe, G. (2019) An Assessment of Human-Wildlife Conflicts in Local Communities Bordering the Western Part of Save Valley Conservancy, Zimbabwe. Global Ecology and Conservation, 20, e00737. https://doi.org/10.1016/j.gecco.2019.e00737
Wilkie, R.D. and Douglas-Hamilton, I. (2018) High-Resolution Tracking Technology Reveals Distinct Patterns in Nocturnal Crop Raiding Behaviour of an African Elephant (Loxodonta africana) in Amboseli, Kenya. Pachyderm, No. 59, 41-48. https://www.savetheelephants.org/wp-content/uploads/2020/02/2018_Wilkie_etal_High-resolutionTrackingTechnologyRevealsDistinctPatternsInNocturnalCropRaidingBehaviourOfEle_8pages.pdf
Songhurst, A., McCulloch, G. and Coulson, T. (2016) Finding Pathways to Human- Elephant Coexistence: A Risky Business. Oryx, 50, 713-720. https://doi.org/10.1017/S0030605315000344
Scheijen, C.P.J., Richards, S.A., Smit, J., Jones, T. and Nowak, K. (2019) Efficacy of Beehive Fences as Barriers to African Elephants: A Case Study in Tanzania. Oryx, 53, 92-99. https://doi.org/10.1017/S0030605317001727
Ndlovu, M., Devereux, E., Chieffe, M., Asklof, K. and Russo, A. (2016) Responses of African Elephants towards a Bee Threat: Its Application in Mitigating Human-Elephant Conflict. South African Journal of Science, 112, 1-5. https://doi.org/10.17159/sajs.2016/20150058
Pozo, R.A., Coulson, T., McCulloch, G., Stronza, A. and Songhurst, A. (2019) Chilli-Briquettes Modify the Temporal Behaviour of Elephants, but Not Their Numbers. Oryx, 53, 100-108. https://doi.org/10.1017/S0030605317001235
Chang’a, A., de Souza, N., Muya, J., Keyyu, J., Mwakatobe, A., Malugu, L., Ndossi, H.P., Konuche, J., Omondi, R., Mpinge, A., Hahn, N., Palminteri, S. and Olson, D. (2016) Scaling-Up the Use of Chili Fences for Reducing Human-Elephant Conflict across Landscapes in Tanzania. Tropical Conservation Science, 9, 921-930. https://doi.org/10.1177/194008291600900220
Gross, E.M., Lahkar, B.P., Subedi, N., Nyirenda, V.R., Lichtenfeld, L.L. and Jakoby, O. (2019) Does Traditional and Advanced Guarding Reduce Crop Losses Due to Wildlife? A Comparative Analysis from Africa and Asia. Journal for Nature Conservation, 50, Article ID: 125712. https://doi.org/10.1016/j.jnc.2019.125712
Evans, L.A. and Adams, W.M. (2016) Fencing Elephants: The Hidden Politics of Wildlife Fencing in Laikipia, Kenya. Land Use Policy, 51, 215-228. https://doi.org/10.1016/j.landusepol.2015.11.008
Hahn, N., Mwakatobe, A., Konuche, J., De Souza, N., Keyyu, J., Goss, M., Chang’a, A., Palminteri, S., Dinerstein, E. and Olson, D. (2017) Unmanned Aerial Vehicles Mitigate Human-Elephant Conflict on the Borders of Tanzanian Parks: A Case Study. Oryx, 51, 513-516. https://doi.org/10.1017/S0030605316000946
Gross, E.M., McRobb, R. and Gross, J. (2016) Cultivating Alternative Crops Reduces Crop Losses Due to African Elephants. Journal of Pest Science, 89, 497-506. https://doi.org/10.1007/s10340-015-0699-2
Somgird, C., Brown, J.L. and Thitaram, C. (2017) Reproductive Control in Elephant: A Tool for Population and Aggression Management. Thai Journal of Veterinary Medicine, 47, 1-6. http://www.thaiscience.info/journals/Article/TJVM/10983993.pdf
Lueders, I., Young, D., Maree, L., Van der Horst, G., Luther, I., Botha, S., Tindall, B., Fosgate, G., Ganswindt, A. and Bertschinger, H.J. (2017) Effects of GnRH Vaccination in Wild and Captive African Elephant Bulls (Loxodonta africana) on Reproductive Organs and Semen Quality. PLoS ONE, 12, e0178270. https://doi.org/10.1371/journal.pone.0178270
Zitzer, H.R. and Boult, V.L. (2018) Vasectomies of Male African Elephants as a Population Management Tool: A Case Study. Bothalia-African Biodiversity & Conservation, 48, a2313. https://doi.org/10.4102/abc.v48i2.2313
Buchholtz, E.K., Redmore, L., Fitzgerald, L.A., Stronza, A., Songhurst, A. and McCulloch, G. (2019) Temporal Partitioning and Overlapping Use of a Shared Natural Resource by People and Elephants. Frontiers in Ecology and Evolution, 7, 117. https://doi.org/10.3389/fevo.2019.00117
Nyirenda, V.R., Nkhata, B.A., Tembo, O. and Siamundele, S. (2018) Elephant Crop Damage: Subsistence Farmers’ Social Vulnerability, Livelihood Sustainability and Elephant Conservation. Sustainability, 10, 3572. https://doi.org/10.3390/su10103572
Noga, S.R., Kolawole, O.D., Thakadu, O.T. and Masunga, G.S. (2017) Claims and Counterclaims: Institutional Arrangements and Farmers’ Response to the Delivery and Adoption of Innovations in the Okavango Delta, Botswana. Journal of Agricultural Education and Extension, 23, 121-139. https://doi.org/10.1080/1389224X.2016.1260485
Evans, L.A. and Adams, W.M. (2018) Elephants as Actors in the Political Ecology of Human-Elephant Conflict. Transactions of the Institute of British Geographers, 43, 630-645. https://doi.org/10.1111/tran.12242
Mumby, H.S. and Plotnik, J.M. (2018) Taking the Elephants’ Perspective: Remembering Elephant Behavior, Cognition and Ecology in Human-Elephant Conflict Mitigation. Frontiers in Ecology and Evolution, 6, 122. https://doi.org/10.3389/fevo.2018.00122
Lindsay, K., Chase, M., Landen, K. and Nowak, K. (2017) The Shared Nature of Africa’s Elephants. Biological Conservation, 215, 260-267. https://doi.org/10.1016/j.biocon.2017.08.021
Selier, S.A.J., Slotow, R. and Di Minin, E. (2016) The Influence of Socioeconomic Factors on the Densities of High-Value Cross-Border Species, the African Elephant. PeerJ, 2016, e2581. https://doi.org/10.7717/peerj.2581
Powell, L.A., Kazahe, U. and Kharuxab, R. (2017) Livestock Farmers Engage in Ecotourism as a Result of Beliefs and Attitudes toward Wildlife on Communal Lands in Namibia. Human Dimensions of Wildlife, 22, 217-230. https://doi.org/10.1080/10871209.2017.1295490
Mayberry, A.L., Hovorka, A.J. and Evans, K.E. (2017) Well-Being Impacts of Human-Elephant Conflict in Khumaga, Botswana: Exploring Visible and Hidden Dimensions. Conservation and Society, 15, 280-291. https://www.jstor.org/stable/26393296
Hacker, C.E. and Miller, L.J. (2016) Zoo Visitor Perceptions, Attitudes, and Conservation Intent after Viewing African Elephants at the San Diego Zoo Safari Park. Zoo Biology, 35, 355-361. https://doi.org/10.1002/zoo.21303
Moolman, L., Ferreira, S.M., Gaylard, A., Zimmerman, D. and Kerley, G.I.H. (2019) The Decline of the Knysna Elephants: Pattern and Hypotheses. South African Journal of Science, 115, 1-7. https://doi.org/10.17159/sajs.2019/4805
Schlossberg, S., Chase, M.J. and Sutcliffe, R. (2019) Evidence of a Growing Elephant Poaching Problem in Botswana. Current Biology, 29, 2222-2228. https://doi.org/10.1016/j.cub.2019.05.061
Chase, M.J., Schlossberg, S., Griffin, C.R., Bouché, P.J.C., Djene, S.W., Elkan, P.W., Ferreira, S., Grossman, F., Kohi, E.M., Landen, K., Omondi, P., Peltier, A., Selier, S.A.J. and Sutcliffe, R. (2016) Continent-Wide Survey Reveals Massive Decline in African Savannah Elephants. PeerJ, 2016, e2354. https://doi.org/10.7717/peerj.2354
Onyango, P.O. and Lesowapir, B. (2016) Demography of a Population with a Long History of Poaching and the Utility of the Individual Identification Technique as a Tool for Monitoring Intermittently Studied Elephant Populations. African Journal of Ecology, 54, 357-364. https://doi.org/10.1111/aje.12297
Rashidi, P., Wang, T., Skidmore, A., Mehdipoor, H., Darvishzadeh, R., Ngene, S., Vrieling, A. and Toxopeus, A.G. (2016) Elephant Poaching Risk Assessed Using Spatial and Non-Spatial Bayesian Models. Ecological Modelling, 338, 60-68. https://doi.org/10.1016/j.ecolmodel.2016.08.002
Beale, C.M., Hauenstein, S., Mduma, S., Frederick, H., Jones, T., Bracebridge, C., Maliti, H., Kija, H. and Kohi, E.M. (2018) Spatial Analysis of Aerial Survey Data Reveals Correlates of Elephant Carcasses within a Heavily Poached Ecosystem. Biological Conservation, 218, 258-267. https://doi.org/10.1016/j.biocon.2017.11.016
Shaffer, M.J. and Bishop, J.A. (2016) Predicting and Preventing Elephant Poaching Incidents through Statistical Analysis, GIS-Based Risk Analysis, and Aerial Surveillance Flight Path Modeling. Tropical Conservation Science, 9, 525-548. https://doi.org/10.1177/194008291600900127
Sibanda, M., Dube, T., Bangamwabo, V.M., Mutanga, O., Shoko, C. and Gumindoga, W. (2016) Understanding the Spatial Distribution of Elephant (Loxodonta africana) Poaching Incidences in the Mid-Zambezi Valley, Zimbabwe Using Geographic Information Systems and Remote Sensing. Geocarto International, 31, 1006-1018. https://doi.org/10.1080/10106049.2015.1094529
Ihwagi, F.W., Wang, T., Wittemyer, G., Skidmore, A.K., Toxopeus, A.G., Ngene, S., King, J., Worden, J., Omondi, P. and Douglas-Hamilton, I. (2015) Using Poaching Levels and Elephant Distribution to Assess the Conservation Efficacy of Private, Communal and Government Land in Northern Kenya. PLoS ONE, 10, e0139079. https://doi.org/10.1371/journal.pone.0139079
Ihwagi, F.W., Skidmore, A.K., Wang, T., Bastille-Rousseau, G., Toxopeus, A.G. and Douglas-Hamilton, I. (2019) Poaching Lowers Elephant Path Tortuosity: Implications for Conservation. Journal of Wildlife Management, 83, 1022-1031. https://doi.org/10.1002/jwmg.21688
Chiyo, P.I., Obanda, V. and Korir, D.K. (2015) Illegal Tusk Harvest and the Decline of Tusk Size in the African Elephant. Ecology and Evolution, 5, 5216-5229. https://doi.org/10.1002/ece3.1769
Raubenheimer, E.J. and Miniggio, H.D. (2016) Ivory Harvesting Pressure on the Genome of the African Elephant: A Phenotypic Shift to Tusklessness. Head and Neck Pathology, 10, 332-335. https://doi.org/10.1007/s12105-016-0704-y
Goldenberg, S.Z., Douglas-Hamilton, I. and Wittemyer, G. (2016) Vertical Transmission of Social Roles Drives Resilience to Poaching in Elephant Networks. Current Biology, 26, 75-79. https://doi.org/10.1016/j.cub.2015.11.005
Pretorius, Y., Garai, M.E. and Bates, L.A. (2019) The Status of African Elephant Loxodonta africana Populations in South Africa. Oryx, 53, 757-763. https://doi.org/10.1017/S0030605317001454
Green, S.E., Davidson, Z., Kaaria, T. and Doncaster, C.P. (2018) Do Wildlife Corridors Link or Extend Habitat? Insights from Elephant Use of a Kenyan Wildlife Corridor. African Journal of Ecology, 56, 860-871. https://doi.org/10.1111/aje.12541
Osipova, L., Okello, M.M., Njumbi, S.J., Ngene, S., Western, D., Hayward, M.W. and Balkenhol, N. (2019) Validating Movement Corridors for African Elephants Predicted from Resistance-Based Landscape Connectivity Models. Landscape Ecology, 34, 865-878. https://doi.org/10.1007/s10980-019-00811-0
Ishida, Y., Van Coeverden de Groot, P.J., Leggett, K.E.A., Putnam, A.S., Fox, V.E., Lai, J., Boag, P.T., Georgiadis, N.J. and Roca, A.L. (2016) Genetic Connectivity across Marginal Habitats: The Elephants of the Namib Desert. Ecology and Evolution, 6, 6189-6201. https://doi.org/10.1002/ece3.2352
Zacarias, D., Bini, L.M. and Loyola, R. (2016) Systematic Review on the Conservation Genetics of African Savannah Elephants. PeerJ, 4, e2567. https://doi.org/10.7717/peerj.2567
Roca, A.L. (2019) African Elephant Genetics: Enigmas and Anomalies. Journal of Genetics, 98, Article No. 83. https://doi.org/10.1007/s12041-019-1125-y
Mondol, S., Moltke, I., Hart, J., Keigwin, M., Brown, L., Stephens, M. and Wasser, S.K. (2015) New Evidence for Hybrid Zones of Forest and Savanna Elephants in Central and West Africa. Molecular Ecology, 24, 6134-6147. https://doi.org/10.1111/mec.13472
Muposhi, V.K., Gandiwa, E., Bartels, P., Makuza, S.M. and Madiri, T.H. (2016) Trophy Hunting and Sustainability: Temporal Dynamics in Trophy Quality and Harvesting Patterns of Wild Herbivores in a Tropical Semi-Arid Savanna Ecosystem. PLoS ONE, 11, e0164429. https://doi.org/10.1371/journal.pone.0164429
Hauenstein, S., Kshatriya, M., Blanc, J., Dormann, C.F. and Beale, C.M. (2019) African Elephant Poaching Rates Correlate with Local Poverty, National Corruption and Global Ivory Price. Nature Communications, 10, Article No. 2242. https://doi.org/10.1038/s41467-019-09993-2
Jung, B.D. (2017) The Tragedy of the Elephants. Wisconsin Law Review, No. 4, 695-737. https://ssrn.com/abstract=3063682
Ogada, D., Botha, A. and Shaw, P. (2016) Ivory Poachers and Poison: Drivers of Africa’s Declining Vulture Populations. Oryx, 50, 593-596. https://doi.org/10.1017/S0030605315001209
Wasser, S.K., Brown, L., Mailand, C., Mondol, S., Clark, W., Laurie, C. and Weir, B.S. (2015) Genetic Assignment of Large Seizures of Elephant Ivory Reveals Africa’s Major Poaching Hotspots. Science, 349, 84-87. https://doi.org/10.1126/science.aaa2457
Kitpipit, T., Thanakiatkrai, P., Penchart, K., Ouithavon, K., Satasook, C. and Linacre, A. (2016) Ivory Species Identification Using Electrophoresis-Based Techniques. Electrophoresis, 37, 3068-3075. https://doi.org/10.1002/elps.201600275
Ziegler, S., Merker, S., Streit, B., Boner, M. and Jacob, D.E. (2016) Towards Understanding Isotope Variability in Elephant Ivory to Establish Isotopic Profiling and Source-Area Determination. Biological Conservation, 197, 154-163. https://doi.org/10.1016/j.biocon.2016.03.008
Seloana, M.Q., Jordaan, J.J., Potgieter, M.J. and Kruger, J.W. (2018) Feeding Patterns of Elephants at the Atherstone Collaborative Nature Reserve. African Journal of Ecology, 56, 445-454. https://doi.org/10.1111/aje.12422
Ward, D., Muller, K. and Shrader, A.M. (2017) Soil Fertility on Granite and Sedimentary Soils Is Associated with Seasonal Differences in Foraging by Elephants. Plant and Soil, 413, 73-81. https://doi.org/10.1007/s11104-016-3067-y
Fullman, T.J., Kiker, G.A., Gaylard, A., Southworth, J., Waylen, P. and Kerley, G.I. (2017) Elephants Respond to Resource Trade-Offs in an Aseasonal System through Daily and Annual Variability in Resource Selection. Koedoe, 59, a1326. https://doi.org/10.4102/koedoe.v59i1.1326
Clegg, B.W. and O’Connor, T.G. (2016) Harvesting and Chewing as Constraints to Forage Consumption by the African Savanna Elephant (Loxodonta africana). PeerJ, 4, e2469. https://doi.org/10.7717/peerj.2469
Kalumanga, E., Mpanduji, D.G. and Cousins, S.A. (2017) Geophagic Termite Mounds as One of the Resources for African Elephants in Ugalla Game Reserve, Western Tanzania. African Journal of Ecology, 55, 91-100. https://doi.org/10.1111/aje.12326
Muposhi, V.K., Gandiwa, E., Chemura, A., Bartels, P., Makuza, S.M. and Madiri, T.H. (2016) Habitat Heterogeneity Variably Influences Habitat Selection by Wild Herbivores in a Semi-Arid Tropical Savanna Ecosystem. PLoS ONE, 11, e0163084. https://doi.org/10.1371/journal.pone.0163084
Stommel, C., Hofer, H., Grobbel, M. and East, M.L. (2016) Large Mammals in Ruaha National Park, Tanzania, Dig for Water When Water Stops Flowing and Water Bacterial Load Increases. Mammalian Biology, 81, 21-30. https://doi.org/10.1016/j.mambio.2015.08.005
Purdon, A. and Van Aarde, R.J. (2017) Water Provisioning in Kruger National Park Alters Elephant Spatial Utilisation Patterns. Journal of Arid Environments, 141, 45-51. https://doi.org/10.1016/j.jaridenv.2017.01.014
Gara, T.W., Wang, T., Skidmore, A.K., Ngene, S.M., Dube, T. and Sibanda, M. (2017) Elephants Move Faster in Small Fragments of Low Productivity in Amboseli Ecosystems: Kenya. Geocarto International, 32, 1243-1253. https://doi.org/10.1080/10106049.2016.1206625
Purdon, A., Mole, M.A., Chase, M.J. and Van Aarde, R.J. (2018) Partial Migration in Savanna Elephant Populations Distributed across Southern Africa. Scientific Reports, 8, Article No. 11331. https://doi.org/10.1038/s41598-018-29724-9
Panagiotopoulou, O., Pataky, T.C., Day, M., Hensman, M.C., Hensman, S., Hutchinson, J.R. and Clemente, C.J. (2016) Foot Pressure Distributions during Walking in African Elephants (Loxodonta africana). Royal Society Open Science, 3, Article ID: 160203. https://doi.org/10.1098/rsos.160203
Rossman, Z.T., Hart, B.L., Greco, B.J., Young, D., Padfield, C., Weidner, L., Gates, J. and Hart, L.A. (2017) When Yawning Occurs in Elephants. Frontiers in Veterinary Science, 4, 22. https://doi.org/10.3389/fvets.2017.00022
Gravett, N., Bhagwandin, A., Sutcliffe, R., Landen, K., Chase, M.J., Lyamin, O.I., Siegel, J.M. and Manger, P.R. (2017) Inactivity/Sleep in Two Wild Free-Roaming African Elephant Matriarchs—Does Large Body Size Make Elephants the Shortest Mammalian Sleepers? PLoS ONE, 12, e0171903. https://doi.org/10.1371/journal.pone.0171903
Wittemyer, G., Keating, L.M., Vollrath, F. and Douglas-Hamilton, I. (2017) Graph Theory Illustrates Spatial and Temporal Features That Structure Elephant Rest Locations and Reflect Risk Perception. Ecography, 40, 598-605. https://doi.org/10.1111/ecog.02379
Tsalyuk, M., Kilian, W., Reineking, B. and Getz, W.M. (2019) Temporal Variation in Resource Selection of African Elephants Follows Long-Term Variability in Resource Availability. Ecological Monographs, 89, e01348. https://doi.org/10.1002/ecm.1348
Wato, Y.A., Prins, H.H., Heitkonig, I., Wahungu, G.M., Ngene, S.M., Njumbi, S. and Van Langevelde, F. (2018) Movement Patterns of African Elephants (Loxodonta africana) in a Semi-Arid Savanna Suggest That They Have Information on the Location of Dispersed Water Sources. Frontiers in Ecology and Evolution, 6, 167. https://doi.org/10.3389/fevo.2018.00167
Goldenberg, S.Z., Douglas-Hamilton, I., Daballen, D. and Wittemyer, G. (2017) Challenges of Using Behavior to Monitor Anthropogenic Impacts on Wildlife: A Case Study on Illegal Killing of African Elephants. Animal Conservation, 20, 215-224. https://doi.org/10.1111/acv.12309
Szott, I.D., Pretorius, Y. and Koyama, N.F. (2019) Behavioural Changes in African Elephants in Response to Wildlife Tourism. Journal of Zoology, 308, 164-174. https://doi.org/10.1111/jzo.12661
Miller, A.K., Hensman, M.C., Hensman, S., Schultz, K., Reid, P., Shore, M., Brown, J., Furthon, K.G. and Lee, S. (2015) African Elephants (Loxodonta africana) Can Detect TNT Using Olfaction: Implications for Biosensor Application. Applied Animal Behaviour Science, 171, 177-183. https://doi.org/10.1016/j.applanim.2015.08.003
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The African Elephant (<i>Loxodonta africana</i>): Mini-Review of an Endangered Species — Oak Academic Publishing