<i>Pakisaurus balochistani</i> (Poripuchia, Slender Titanosauria, Sauropoda) Associated Skeletons Found from the Latest Maastrichtian Vitakri Formation of Pakistan and Referred Fossils from India; Filling of Important Missing Links of <i>Isisaurus colberti</i> Titanosaur Found from Pakistan — Oak Academic Publishing
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<i>Pakisaurus balochistani</i> (Poripuchia, Slender Titanosauria, Sauropoda) Associated Skeletons Found from the Latest Maastrichtian Vitakri Formation of Pakistan and Referred Fossils from India; Filling of Important Missing Links of <i>Isisaurus colberti</i> Titanosaur Found from Pakistan
Geological Survey of Pakistan, Muzaffarabad, Pakistan
1 Geological Survey of Pakistan, Muzaffarabad, Pakistan
Recently two slender and medium to large sized titanosaur taxa like Pakisaurus balochistani and Isisaurus colberti of pakisaurids Poripuch along with small sized stocky Saraikimasoom and medium sized stocky Gspsaurus of gspsaurids Poripuch have been recognized from Indo-Pakistan subcontinent. The diagnosis and description of Pakisaurus balochistani and some missing elements of Isisaurus colberti discovered from Pakistan are added here for understanding of possible fuller anatomy of slender titanosaurs. Further so far no any research information regarding the comparison of titanosaurs of India and Pakistan is available. Here present research provides the comparison between titanosaurs recognized from Pakistan and India.
Sahni, A. (2001) Dinosaurs of India. National Book Trust, Delhi, 110.
Malkani, M.S. and Anwar, C.M. (2000) Discovery of First Dinosaur Fossil in Pakistan, Barkhan District, Balochistan. Geological Survey of Pakistan Information Release, 732, 1-16.
Malkani, M.S. (2006) Lithofacies and Lateral Extension of Latest Cretaceous Dinosaur Beds from Sulaiman Foldbelt, Pakistan. Sindh University Research Journal (Science Series), 38, 1-32.
Malkani, M.S. (2019) Cretaceous Stratigraphy of Pakistan. Open Journal of Geology, 9, 671-673. https://doi.org/10.4236/ojg.2019.910071
Malkani, M.S. (2003) Discovery of Partial Skull and Dentary of Titanosauria (Sauropod Dinosaur) from the Late Cretaceous Pab Formation of Vitakri Area, Barkhan District, Balochistan, Pakistan. Journal of Himalayan Earth Sciences, 36, 65-71.
Malkani, M.S. (2003) Pakistani Titanosauria; Are Armoured Dinosaurs? Journal of Himalayan Earth Sciences, 36, 85-91.
Malkani, M.S. (2003) First Jurassic Dinosaur Fossils Found from Kirthar Range, Khuzdar District, Balochistan, Pakistan. Journal of Himalayan Earth Sciences, 36, 73-83.
Malkani, M.S. (2004) Saurischian Dinosaurs from Late Cretaceous of Pakistan. In: Hussain, S.S. and Akbar, H.D., Eds., Proceedings of Fifth Pakistan Geological Congress, National Geological Society of Pakistan, Pakistan Museum of Natural History, Islamabad, 14-15 April 2004, 71-73.
Malkani, M.S. (2006) Biodiversity of Saurischian Dinosaurs from the Latest Cretaceous Park of Pakistan. Journal of Applied and Emerging Sciences, 1, 108-140.
Malkani, M.S. (2006) Cervicodorsal, Dorsal and Sacral Vertebrae of Titanosauria (Sauropod Dinosaurs) Discovered from the Latest Cretaceous Dinosaur Beds/Vitakri Member of Pab Formation, Sulaiman Foldbelt, Pakistan. Journal of Applied and Emerging Sciences, 1, 188-196.
Malkani, M.S. (2008) Marisaurus (Balochisauridae, Titanosauria) Remains from the Latest Cretaceous of Pakistan. Sindh University Research Journal (Science Series), 40, 55-78.
Malkani, M.S. (2008) First Articulated Atlas-Axis Complex of Titanosauria (Sauropoda, Dinosauria) Uncovered from the Latest Cretaceous Vitakri Member (Dinosaur Beds) of Upper Pab Formation, Kinwa Locality of Sulaiman Basin, Pakistan. Sindh University Research Journal (Science Series), 40, 55-70.
Malkani, M.S. (2010) Osteoderms of Pakisauridae and Balochisauridae (Titanosauria, Sauropoda, Dinosauria) in Pakistan. Journal of Earth Science, 21, 198-203. https://doi.org/10.1007/s12583-010-0212-z
Malkani, M.S. (2009) New Balochisaurus (Balochisauridae, Titanosauria, Sauropoda) and Vitakridrinda (Theropoda) Remains from Pakistan. Sindh University Research Journal (Science Series), 41, 65-92.
Malkani, M.S. (2010) New Pakisaurus (Pakisauridae, Titanosauria, Sauropoda) Remains and Cretaceous Tertiary (K-T) Boundary from Pakistan. Sindh University Research Journal (Science Series), 42, 39-64.
Wilson, J.A., Barrett, P.M. and Carrano, M.T. (2011) An Associated Partial Skeleton of Jainosaurus cf. septentrionalis (Dinosauria: Sauropoda) from the Late Cretaceous of Chhota Simla, Central India. Palaeontology, 54, 981-998. https://doi.org/10.1111/j.1475-4983.2011.01087.x
Malkani, M.S. (2014) Titanosaurian Sauropod Dinosaurs from the Latest Cretaceous of Pakistan. Abstract Volume of 2nd symposium of IGCP 608 “Cretaceous Ecosystem of Asia and Pacific”, Tokyo, Japan, 4-6 September, 108-111.
Malkani, M.S. (2015) Dinosaurs, Mesoeucrocodiles, Pterosaurs, New Fauna and Flora from Pakistan. Geological Survey of Pakistan, Information Release, 823, 1-32.
Malkani, M.S. (2015) Titanosaurian Sauropod Dinosaurs from Pakistan. In: Zhang, Y., Wu, S.Z. and Sun, G. Eds., Abstract Volume, 12th Symposium on “Mesozoic Terrestrial Ecosystems (MTE 12), and 3rd Symposium of International Geoscience Program (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific”, Paleontological Museum of Liaoning/Shenyang Normal University, Shenyang, China, 15-20 August, 2015, 93-98.
Malkani, M.S. (2019) Medium Sized Stocky Titanosaur from South Asia. Open Journal of Geology, 9, 631-634. https://doi.org/10.4236/ojg.2019.910060
Malkani, M.S. (2020) First Skull of Medium Sized Titanosaur from Indo-Pakistan Subcontinent Found from Found from the Latest Maastrichtian Vitakri Formation of Pakistan; Associated Cranial and Postcranial Skeletons of Gspsaurus pakistani (Poripuchia, Stocky Titanosauria, Sauropoda) from Pakistan and India. Open Journal of Geology, 10, (In Press). https://doi.org/10.4236/ojg.2019.910059
Malkani, M.S. (2019) Smallest Titanosaur from Indo-Pakistan Landmass. Open Journal of Geology, 9, 627-630. https://doi.org/10.4236/ojg.2019.910059
Malkani, M.S. (2020) First Snout with Complete Teeth Row of Titanosaur from Indo-Pakistan Subcontinent Found from the Latest Maastrichtian Vitakri Formation of Pakistan; Associated Cranial and Postcranial Skeletons of Saraikimasoom vitakri (Poripuchia, Stocky Titanosauria, Sauropoda) from Pakistan and Referred Fossils from India. Open Journal of Geology, 10, (In Press).
Malkani, M.S. (2019) Large Titanosaur from Indo-Pakistan Peninsula. Open Journal of Geology, 9, 635-638. https://doi.org/10.4236/ojg.2019.910061
Jain, S.L. and Bandyopadhyay, S. (1997) New Titanosaurid (Dinosauria: Sauropoda) from the Late Cretaceous of Central India. Journal of Vertebrate Paleontology, 17, 114-136. https://doi.org/10.1080/02724634.1997.10010958
Malkani, M.S., Islam, N., Mahmood, Z. and Rajper, R.H. (2017) New Celestite Deposits from Laki Shale and Limestone, and Some Evidences of Dinosaurs and Associated Vertebrates from Laki Anticline, Sindh Province, Pakistan. Geological Survey of Pakistan, Information Release, 1017, 1-18.
Malkani, M.S. (2019) Recently Discovered Basilosaurid, Baluchithere Rhinoceros, Horses, Sea Cow, Proboscidean, Eucrocodile, Pterosaurs, Plesiosaur, Fishes, Invertebrates and Wood Fossils, Tracks and Trackways of Dinosaurs from Pakistan; Comparison of Recognized Four Titanosaur Taxa of Indo-Pakistan with Madagascar. Open Journal of Geology, 9, 919-955. https://doi.org/10.4236/ojg.2019.912098
Owen, R. (1842) Report on British Fossil Reptiles, Pt. II. Reptiles. Report of the British Association for the Advancement of Science, No. 1841, 60-204.
Seeley, H.G. (1888) The Classification of the Dinosauria. Report of the British Association for the Advancement of Science, No. 1887, 698-699. https://doi.org/10.1017/S0016756800156006
Marsh, O.C. (1878) Principal Characters of American Jurassic Dinosaurs. Part I. American Journal of Science (Series 3), 16, 411-416. https://doi.org/10.2475/ajs.s3-16.95.411
Bonaparte, J.F. and Coria, R.A. (1993) A New and Huge Titanosaur Sauropod from the Rio Limay Formation (Albian-Cenomanian) of Neuquen Province, Argentina. Ameghiniana, 30, 271-282.
Swinton, W.E. (1947) New Discoveries of Titanosaurus indicus Lyd. Annals and Magazine of Natural History (Series 11), 14, 112-123. https://doi.org/10.1080/00222934708654616
Mathur, U.B. and Srivastava, S. (1987) Dinosaur Teeth from Lameta Group (Upper Cretaceous) of Kheda District, Gujurat. Journal of the Geological Society of India, 29, 554-566.
Wilson, J.A. and Upchurch, P. (2003) A Revision of Titanosaurus Lydekker (Dinosauria, Sauropoda), the First Dinosaur Genus with a ‘Gondwanan’ Distribution. Journal of Systematic Paleontology, 1, 125-160. https://doi.org/10.1017/S1477201903001044
Huene, F.V. and Matley, C.A. (1933) Cretaceous Saurischia and Ornithischia of the Central Provinces of India. Paleontologia Indica, 21, 1-74.
Wilson, J.A. and Mohabey, D.M. (2006) A Titanosauriform (Dinosauria: Sauropoda) Axis from the Lameta Formation (Upper Cretaceous: Maastrichtian) of Nand, Central India. Journal of Vertebrate Paleontology, 26, 471-479. https://doi.org/10.1671/0272-4634(2006)26[471:ATDSAF]2.0.CO;2
Lydekker, R. (1877) Notes of New and Other Vertebrata from Indian Tertiary and Secondary Rocks. Record Geological Survey of India, 10, 30-43.
Malkani, M.S. (2015) Mesozoic Stratigraphy of Pakistan: Special Emphasis on Indus Basin’s Revised Stratigraphy. In: Zhang, Y., Wu, S.Z. and Sun, G., Eds., Abstract Volume, 12th Symposium on “Mesozoic Terrestrial Ecosystems (MTE 12), and 3rd Symposium of International Geoscience Program (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific”, Paleontological Museum of Liaoning/Shenyang Normal University, Shenyang, 15-20 August 2015, 255-260.
Malkani, M.S. (2014) Records of Fauna and Flora from Pakistan; Evolution of Indo-Pakistan Peninsula. Abstract Volume of 2nd Symposium of International Geoscience Program (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific”, Tokyo, Japan, 4-6 September 2014, 165-168.
Vredenburg, E.W. (1908) The Cretaceous Orbitoides of India. IBID, 36, 171-213.
Williams, M.D. (1959) Stratigraphy of the Lower Indus Basin, West Pakistan. 5th World Petroleum Congress, New York, 277-394.
Jones, A.G., Manistere, B.E., Oliver, R.L., Willson, G.S. and Scott, H.S. (1961) Reconnaissance Geology of Part of West Pakistan (Colombo Plan Co-Operative Project Conducted and Compiled by Hunting Survey Corporation). Government of Canada, Toronto, 550 p.
Malkani, M.S. and Mahmood, Z. (2017) Stratigraphy of Pakistan. Geological Survey of Pakistan, Memoir Volume, 24, 1-134.
Malkani, M.S. and Mahmood, Z. (2016) Revised Stratigraphy of Pakistan. Geological Survey of Pakistan, Record Volume, 127, 1-87.
Prasad, G.V.R. (1989) Vertebrate Fauna from the Infra and Intertrappean Beds of Andhra Pradesh: Age Implications. Journal of the Geological Society of India, 34, 161-173.
Sahni, A. and Bajpai, S. (1991) Eurasiatic Animals in the Upper Cretaceous Nonmarine Biotas of Peninsular India. Cretaceous Research, 12, 177-183. https://doi.org/10.1016/S0195-6671(05)80023-0
Sanz, J.L., Powell, J.E., LeLoeuff, J., Martinez, R. and Pereda-Suberiola, X. (1999) Sauropod Remains from the Upper Cretaceous of Lano (Northcentral Spain), Titanosaur Phylogenetic Relationships. Estudios del Museo de Ciencias Naturales de Nava, 14, 235-255.
Powell, J.E. (1990) Epachthosaurus sciutoi g et sp. nov., un Nuevo dinosaurio titanosaurido del Cretacico de Patagomia. Actas V Congreso Argentino de Paleontologia y Bioestratigrafia, 1, 123-128.
Mannion, P.D., Upchurch, P., Barnes, R.N. and Mateus, O. (2013) Osteology of the Late Jurassic Portuguese Sauropod Dinosaur Lusotitan atalaiensis (Macronaria) and the Evolutionary History of Basal Titanosauriformes. Zoological Journal of the Linnean Society, 168, 98-206. https://doi.org/10.1111/zoj.12029
McIntosh, J.S. (1990) Sauropoda. In: Weishampel, D.B., Dodson, P. and ósmolska, H., Eds., The Dinosauria, University California Press, Berkeley, CA, 345-401.
Jacobs, L.L., Winkler, D.A., Downs, W.R. and Gomani, E.M. (1993) New material of an Early Cretaceous Titanosaurid Sauropod Dinosaur from Malawi. Palaeontology, 36, 523-534.
Upchurch, P. (1995) The Evolutionary History of Sauropod Dinosaurs. Philosophical Transactions of the Royal Society of London, Series B, 349, 365-390. https://doi.org/10.1098/rstb.1995.0125
Upchurch, P. (1998) The Phylogenetic Relationships of Sauropod Dinosaurs. Zoological Journal of the Linnean Society, 124, 43-103. https://doi.org/10.1111/j.1096-3642.1998.tb00569.x
Salgado, L., Coria, R.A. and Calvo, J.O. (1997) Evolution of Titanosaurids Sauropods. I: Phylogenetic Analysis Based on the Postcranial Evidence. Ameghiniana, 34, 3-32.
Wilson, J.A. (2002) Sauropod Dinosaur Phylogeny: Critique and Cladistic Analysis. Zoological Journal of the Linnean Society, 136, 217-276. https://doi.org/10.1046/j.1096-3642.2002.00029.x
Mannion, P.D. and Calvo, J.O. (2011) Anatomy of the Basal Titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the Mid-Cretaceous (Albian-early Cenomanian) Río Limay Formation, Neuquén Province, Argentina: Implications for Titanosaur Systematics. Zoological Journal of the Linnean Society, 163, 155-181. https://doi.org/10.1111/j.1096-3642.2011.00699.x
Canudo, J.I., Royo-Torres, R. and Cuenca-Bescós, G. (2008) A New Sauropod: Tastavinsaurus sanzi gen. et sp. nov. from the Early Cretaceous (Aptian) of Spain. Journal of Vertebrate Paleontology, 28, 712-731. https://doi.org/10.1671/0272-4634(2008)28[712:ANSTSG]2.0.CO;2
Wang, X., You, H., Meng, Q., Gao, C., Cheng, X. and Liu, J. (2007) Dongbeititan dongi, the First Sauropod Dinosaur from the Lower Cretaceous Jehol Group of Western Liaoning Province, China. Acta Geologica Sinica, 81, 911-916. https://doi.org/10.1111/j.1755-6724.2007.tb01013.x
Dong, Z. (1990) Sauropoda from the Kelameili Region of the Junggar Basin, Xinjiang Autonomous Region. Vertebrata PalAsiatica, 28, 43-58.
Sekiya, T. (2011) Re-Examination of Chuanjiesaurus anaensis (Dinosauria: Sauropoda) from the Middle Jurassic Chuanjie Formation, Lufeng County, Yunnan Province, Southwest China. Memoir of Fukui Prefectural Dinosaur Museum, 10, 1-54.
Bonaparte, J.F., Heinrich, W.-D. and Wild, R. (2000) Review of Janenschia Wild, with the Description of a New Sauropod from the Tendaguru Beds of Tanzania and a Discussion on the Systematic Value of Procoelous Caudal Vertebrae in the Sauropoda. Palaeontographica, Abteilung A, 256, 25-76.
Casanovas, M.L., Santafe, J.V. and Sanz, J.L. (2001) Losillasaurus giganteus, un nuevo sauropodo del transito Jurasico-Cretacico de la cuenca de ‘Los Serranos’ (Valencia, Espana). Paleontologia i Evolucio, 32-33, 99-122.
Janensch, W. (1950) Die wirbelsaule von Brachiosaurus brancai. Palaeontographica, 3, 27-93.
D’Emic, M.D. (2012) The Early Evolution of Titanosauriform Sauropod Dinosaurs. Zoological Journal of the Linnean Society, 166, 624-671. https://doi.org/10.1111/j.1096-3642.2012.00853.x
D’Emic, M.D. (2013) Revision of the Sauropod Dinosaurs of the Early Cretaceous Trinity Group, Southern USA, with the Description of a New Genus. Journal of Systematic Palaeontology, 11, 707-726. https://doi.org/10.1080/14772019.2012.667446
Taylor, M.P., Wedel, M.J. and Cifelli, R.L. (2011) A New Sauropod Dinosaur from the Lower Cretaceous Cedar Mountain Formation, Utah, USA. Acta Palaeontologica Polonica, 56, 75-98. https://doi.org/10.4202/app.2010.0073
You, H.-L., Tang, F. and Luo, Z. (2003) A New Basal Titanosaur (Dinosauria: Sauropoda) from the Early Cretaceous of China. Acta Geologica Sinica, 77, 424-429. https://doi.org/10.1111/j.1755-6724.2003.tb00123.x
Gonzalez Riga, B.J. (2003) A New Titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza Province, Argentina. Ameghiniana, 40, 155-172.
Royo-Torres, R. (2009) El saurópodo de Penarroya de Tastavins. Instituto de Estudios Turolenses-Fundación Conjunto Paleontológico de Teruel-Dinópolis, Monografías Turolenses, 6, 1-548.
Curry Rogers, K.A. and Forster, C.A. (2004) The Skull of Rapetosaurus krausei (Sauropoda: Titanosauria) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology, 24,121-144. https://doi.org/10.1671/A1109-10
Yadagiri, P. and Ayyasami, K. (1987) A Carnosaurian Dinosaur from the Kallamedu Formation (Maestrichtian Horizon), Tamil Nadu. Geological Survey of India Special Publications, 1, 523-528.
Malkani, M.S. (2007) Trackways Evidence of Sauropod Dinosaurs Confronted by a Theropod Found from Mid Jur. Samana Suk Limestone of Pakistan. Sindh University Research Journal (Science Series), 39, 1-14.
Nizami, A.R. and Fahim, M. (2015) The Discovery of Ichno-Fossils of the Middle Jurassic Dinosaurs in the Central Salt Range, District Khoshab, Punjab, Sub-Himalayas, Pakistan. Punjab University Journal of Zoology, 30, 37-41.
Malkani, M.S. (2015) Footprints and Trackways of Dinosaurs from Indo-Pakistan Subcontinent-Recent Advances in Discoveries from Pakistan. In: Zhang, Y., Wu, S.Z. and Sun, G., Eds., Abstract Volume, 12th Symposium on “Mesozoic Terrestrial Ecosystems (MTE 12), and 3rd Symposium of International Geoscience Program (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific”, Paleontological Museum of Liaoning/Shenyang Normal University, Shenyang, 15-20 August 2015, 186-191.
Hunt, A.P., Lockley, M.G., Lucas, S.G. and Meyer, C.A. (1994) The Global Sauropod Fossil Record. GAIA, 10, 261-279.
Wilson, J.A., Malkani, M.S. and Gingerich, P.D. (2005) A Sauropod Braincase from the Pab Formation (Upper Cretaceous, Maastrichtian) of Balochistan, Pakistan. Gondwana Geological Magazine, Special Volume, 8, 101-109.
Malkani, M.S. (2017) A Review of Pakisaurus titanosaur: Associated Skeletons of Sauropod and Theropod Dinosaurs and Mesoeucrocodiles from Pakistan: Discussion on Titanosaurs of Indo-Pak Subcontinent. In: Cheong, D., Lee, Y. and Kim, D., Eds., Commemoration of the 70th Anniversary of the Geological Society of Korea, Proceedings and Fieldguide book for the Fifth International Symposium of International Geoscience Programme IGCP Project 608 “Cretaceous Ecosystems and Their Responses to Paleoenvironmental Changes in Asia and the Western Pacific”, Jeju Island, Korea, 22-28 October 2017, 116-119.
Malkani, M.S. (2020) Exploration of Latest Theropods, Mesoeucrocodiles and Pterosaurs from Pakistan. (In Process)
Malkani, M.S. (2019) Vitakrisaurus saraiki Theropod from South Asia. Open Journal of Geology, 9, 643-645. https://doi.org/10.4236/ojg.2019.910063
Malkani, M.S. (2019) Induszalim bala Mesoeucrocodile from Pakistan. Open Journal of Geology, 9, 623-626. https://doi.org/10.4236/ojg.2019.910058
Wilson, J.A., Malkani, M.S. and Gingerich, P.D. (2001) New Crocodyliform (Reptilia, Mesoeucrocodylia) form the Upper Cretaceous Pab Formation of Vitakri, Balochistan (Pakistan). Contributions from Museum of Paleontology, University of Michigan, 30, 321-336.
Malkani, M.S. (2010) Updated Stratigraphy and Mineral potential of Sulaiman (Mid. Indus) Basin, Pakistan. Sindh University Research Journal (Science Series), 42, 39-66.
Dubey, V.S. and Narain, K. (1946) A Note on the Occurrence of Pterosauria in India. Current Science, 15, 287-288. https://doi.org/10.1525/as.1946.15.18.01p0090d
Malkani, M.S. and Mahmood, Z. (2017) Mineral Resources of Pakistan: Provinces and Basins Wise. Geological Survey of Pakistan, Memoir Volume, 25, 1-179.
Malkani, M.S. and Mahmood, Z. (2016) Mineral Resources of Pakistan: A Review. Geological Survey of Pakistan, Record Volume, 128, 1-90.
Malkani, M.S. (2011) Stratigraphy, Mineral Potential, Geological History and Paleobiogeography of Balochistan Province, Pakistan. Sindh University Research Journal (Science Series), 43, 269-290.
Malkani, M.S. (2015) Geodiverse and Biodiverse Heritage of Pakistan Demands for Protection as National and Global Geoparks: An Innovation for the Sustainable Development of Pakistan. In: Zhang, Y., Wu, S.Z. and Sun, G., Eds., Abstract Volume, 12th Symposium on “Mesozoic Terrestrial Ecosystems (MTE 12) and 3rd Symposium of International Geoscience Program (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific”, Paleontological Museum of Liaoning/Shenyang Normal University, Shenyang, China, 15-20 August 2015, 247-249.
Wilson, J.A., Mohabey, D.M., Lakra, P. and Bhadran, A. (2019) Titanosaur (Dinosauria, Sauropoda) Vertebrae from the Upper Cretaceous Lameta Formation of Western and Central India. Contributions from the Museum of Paleontology, University of Michigan, 33, 1-27.